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The Penguins of Madagascar is a spin-off of the Madagascar films. The series follows the adventures of the four penguin protagonists: Skipper, Kowalski, Rico, and Private, who perform various paramilitary-like missions to protect their home in the Central Park Zoo. The penguins often have to deal with problems caused, or made worse, by King Julien XIII (a ring-tailed lemur), Maurice (an aye-aye), and Mort (a mouse lemur). It is not known how the penguins and lemurs arrived at the zoo after their adventures in both Madagascar films, although the penguins and chimpanzees are seen flying away in a plane at the end of Madagascar 2. During the show's opening title, the penguins are also seen opening a crate that reads "Madagascar" and contains the three lemur characters.
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Cast:
| King Julien XIII (Danny Jacobs) | normally shortened to King Julien, is a fun loving, narcissistic ring-tailed lemur |
| Kowalski (Jeff Bennett) | acts as the group strategist and gadgeteer |
| Marlene (Nicole Sullivan) | a female otter who was transferred to the Central Park Zoo from an aquarium in California |
| Maurice (Kevin Michael Richardson) | an aye-aye, is one of King Julien's subjects |
| Mort (Andy Richter) | an excitable, dimwitted, accident-prone mouse lemur |
| Private (James Patrick Stuart) | is the emotionally sensitive rookie of the group |
| Rico (John DiMaggio) | the team's weapons and explosives specialist, who mainly communicates through grunts and squeals |
| Skipper (Tom McGrath) | the leader of the penguins; he devises tactics and gives orders |